Tel: +47 79 02 33 00 | post@unis.no
Steve Coulson
Position: Professor, Terrestrial ecology
Phone:

+47 79 02 33 34

Cellular Phone: +47
E-mail: steve.coulson<a>unis.no
Webpage: Personal web page
Room:  
Courses: AB-201;  AB-203; AB-206,
UNIS contact: AB-325, AB-825, AB-333, AB-833

 

The terrestrial and freshwater invertebrate biodiversity of the archipelagoes of the Barents Sea; Svalbard, Franz Josef Land and Novaya Zemlya Soil Biology and Biochemistry 2014, 68; 440-470.

(Coulson, S.J., Convey, P., Aakra, K., Aarvik, L., Ávila-Jiménez, M.L., Babenko, A., Biersma, E., Boström, S., Brittain, J., Carlsson, A.M., Christoffersen, K.S., De Smet, W.H., Ekrem, T., Fjellberg, A., Füreder, L., Gustafsson, D., Gwiazdowicz, D.J.,  Holmstrup M., Hansen, L.O., Holmnstrup, M., Kaczmarek, L.,  Kolicka, M., Kuklin, V., Lakka, H-K., Lebedeva, N., Makarova, O., Maraldo, K., Melekhina, E., Ødegaard, F., Pilskog, H.E., Simon, J.C., Sohlenius, B., Solhøy, T., Søli, G.,  Stur, E., Tanaevitch, A., Taskaeva, A., Velle, G. Zawierucha, K. and Zmudczyńska-Skarbek, K.)

Short CV  

 

  POSTER - Invertebrate biodiversity of the Archipelagoes of the Barents Sea  
  POSTER - Survival of low temperatures by Arctic soil invertebrates  
  Checklist of the Terrestrial and freshwater invertebrate fauna of Svalbard (Pdf)  
  SPIDER (Svalbard Pictographic Invertebrate Database and Educational Resource)  
  AVIFauna project "On borrowed wings"- written by Media Research  
  AVIFaauna project "Flying without wings"- written by Media Research  
  What is SPIDER?  
  Arctic Bug Group (external page under construction)  
  AVIFauna webpage and blog (external page under constrution)  

 

 

Steve Coulson in UNIS news:

 

  Mice on ice!                                                                                                                  (06.09.2012)  
  The snow, the aphid and climate change                                                            (11.07.2012)  
  Insects in Svalbard; what is there and what do we know?                            (18.06.2012)  
  Introducing Isohypsibius coulsoni                                                                       (12.03.2012)  
A long journey for terrestrial invertebrates                                                      (20.09.2011)
Small, but far from insignificant                                                                           (13.07.2011)
Norwegian-Russian workshop on invertebrates                                               (25.03.2011)
Creepy crawlies on Svalbard; unseen and often forgotten                            (01.08.2010)
The Svalbard flora and invertebrate fauna                                                        (29.06.2010)  
  Creepy-crawlies in a seabird’s nest                                                                      (09.07.2009)  

 

Publications

In press

  • Elsterová J., Černý J., Müllerová J., Šíma R., Coulson S.J., Lorentzen E., Strøm H., Grubhoffer L. Search for tick-borne pathogens in the Svalbard archipelago and Jan Mayen.  Polar Research. 
  • Coulson S.J. The alien terrestrial invertebrate fauna of the High Arctic archipelago of Svalbard: potential implications for the native flora and fauna. Polar Research
  • Convey, P., Abbandonato, H.D.A., Bergan, F., Beumer, L.T., Biersma, E.M., Bråthen, V.S., D'Imperio, L., Jensen, C.K., Nilsen, S.,  Paquin, K., Stenkewitz, U., Svoen, M.E., Winkler, J., Müller, E. and Coulson, S.J. (2014). Survival of rapidly fluctuating natural low winter temperatures by Arctic soil invertebrates. Manuscript resulting from project on UNIS masters/Ph.D. Arctic Winter Ecology course (AB329/829) 2012-13 Journal of Thermal Biology DOI: 10.1016/j.jtherbio.2014.07.009

 Published

2015

  • Coulson S.J., Fjellberg A., Melekhina E.N.,  Taskaeva A.A:, Lebedeva N.V., Belkina O., Seniczak S., Seniczak A. and Gwiazdowicz D.J. Microarthropod communities of disturbed or imported soils in the High Arctic; the abandoned coal mining town of Pyramiden, Svalbard. Biodiversity and Conservation DOI 10.1007/s10531-015-0885-9

2014

  • Hansen B.B., Isaksen K.,  Benestad R.E., Kohler J., Pedersen Å.Ø.,  Loe L.E.,  Coulson S.J., Larsen J.O. and Varpe Ø. Warmer, Wetter, Wilder Winters: Characteristics and implications of an extreme weather event in the High Arctic. Environmental Research Letters 9 (2014) 114021 (10pp) doi:10.1088/1748-9326/9/11/114021
  • Dabert M., Coulson S.J., Gwiazdowicz D.J., Moe B., Hanssen S.A., Biersma E.M., Pilskog H.E. and Dabert J. Different modes of speciation in feather mites (Analgoidea) inhabiting the same host: The case of Zachvatkinia and Alloptes living on arctic and long-tailed skuas. Experimental and Applied Acarology doi 10.1007/s10493-014-9856-1
  • Seniczak, S., Seniczak, A., Gwiazdowicz, D.J. & Coulson S.J. (2014). Oribatid and gamasid mites (Acari) of chosen patches of moss-grass tundra in Svalbard (Spitsbergen, Norway). Arctic, Antarctic, and Alpine Research, 46; 591–599.
  • Coulson, S.J., Convey, P., Aakra, K., Aarvik, L., Ávila-Jiménez, M.L., Babenko, A., Biersma, E., Boström, S., Brittain, J., Carlsson, A.M., Christoffersen, K.S., De Smet, W.H., Ekrem, T., Fjellberg, A., Füreder, L., Gustafsson, D., Gwiazdowicz, D.J.,  Hansen, L.O., Holmstrup M., Kaczmarek, L.,  Kolicka, M., Kuklin, V., Lakka, H-K., Lebedeva, N., Makarova, O., Maraldo, K., Melekhina, E., Ødegaard, F., Pilskog, H.E., Simon, J.C., Sohlenius, B., Solhøy, T., Søli, G.,  Stur, E., Tanaevitch, A., Taskaeva, A., Velle, G. Zawierucha, K. and Zmudczyńska-Skarbek, K. (2014). The terrestrial and freshwater invertebrate biodiversity of the archipelagoes of the Barents Sea; Svalbard, Franz Josef Land and Novaya Zemlya. Soil Biology and Biochemistry 68; 440-470.
  • Pilskog, H.E., Solhøy, T., Gwiazdowicz, D.J., Grytnes, J.A. and Coulson S.J. (2014). Invertebrates in the nests of migrating passerine, wild fowl and sea birds breeding in the High Arctic, Svalbard. Polar Biology 37:981–998. DOI 10.1007/s00300-014-1495-9 
  • Coulson S.J., Schatz H., Gwiazdowicz D.J. and Solhøy T. (2014). The oribatid and mesostigmatid mite fauna (Acari) of the High Arctic island of Hopen. Polish Polar Research. 35; 133–139, 2014 doi: 10.2478/popore−2014−0002 
  • Teodorowicz E., Gwiazdowicz D.J. & Coulson S.J (2014). Redescription of Antennoseius (Vitzthumia) oudemansi (Acari, Mesostigmata) from Spitsbergen. Entomologica Fennica. 25; 27–42.

2013

  • Coulson S.J.  (2013) The terrestrial invertebrate fauna of the Svalbard archipelago in a changing world; history of research and challenges. Canadian Entomologist 145; 131-146. doi: 10.4039/tce.2012.110
  • Zawierucha, K., Coulson S.J., Michalczyk L. and Kaczmarek. L. (2013) Current knowledge on the Tardigrada of Svalbard with the first records of water bears from Nordaustlandet (High Arctic). Polar Research 2013, 32, 20886, http://dx.doi.org/10.3402/polar.v32i0.20886 
  • Coulson, S.J.,  Fjellberg, A., Gwiazdowicz, D.J., Lebedeva,  N.V.,  Melekhina, E.N., Solhøy, T.,  Erséus, C., Maraldo, K., Miko, L., Schatz,  H., Schmelz, R.M., Søli, G. and Stur, E. (2013) Introduction of invertebrates into the High Arctic via imported soils: the case of Barentsburg in Svalbard. Biological Invasions 15;1–5. doi 10.1007/s10530-012-0277-y  
  • Coulson, S.J.,  Fjellberg, A., Gwiazdowicz, D.J., Lebedeva,  N.V.,  Melekhina, E.N., Solhøy, T.,  Erséus, C., Maraldo, K., Miko, L., Schatz,  H., Schmelz, R.M., Søli, G. and Stur, E. (2013). The invertebrate fauna of anthropogenic soils in the High Arctic settlement of Barentsburg; Svalbard. Polar Research 32, 19273, http://dx.doi.org/10.3402/polar.v32i0.19273
  • Carlsson, A.M., Irvine, R.J., Wilson, K.  and Coulson, S.J. (2013). Adaptations to the Arctic: low-temperature development and cold tolerance in the free-living stages of a parasitic nematode from Svalbard. Polar Biology 36; 997-1005 doi: 10.1007/s00300-013-1323-7

2012

  • Convey, P., Aitken, S., di Prisco, G., Gill, M.J., Coulson, S.J., Barry, T., Jónsdóttir, I.S., Dang, P.Y., Hik, D., Kulkarni T., & Lewis, G. (2012). The impacts of climate change on circumpolar biodiversity Biodiversity (Key Note paper Special Issue: Circumpolar Biodiversity) 13;134-143 doi: 10.1080/14888386.2012.732556
  • Coulson S.J. & Midgley N.G. (2012) The role of glacier mice in the invertebrate colonisation of glacial surfaces: the moss balls of the Falljökull, Iceland. Polar Biology 35;1651–1658. doi: 10.1007/s00300-012-1205-4
  • Gwiazdowicz D.J., Solhøy T., Coulson S.J., Lebedeva N. & Melekhina E. (2012). Vulgarogamasus immanis (Acari; Mesostigmata) in Svalbard. Polish Polar Research 33; 35-39.
  • Carlsson, A.M., Wilson, K., Irvine, R.J., Piertney, S.B., Halvorsen, O., Coulson, S.J., Stien, A. & Albon S.D. (2012) Disease transmission in an extreme environment: nematode parasites infect reindeer during the Arctic winter. International Journal of Parasitology 42; 789-795.
  • Gwiazdowicz D.J., Coulson S.J., Grytnes J.A. & Pilskog H.E. (2012) Dermanyssus hirundinis (Acari, Mesostigmata) new parasitic mite to Spitsbergen, Svalbard. Acta Parasitologica 57(4), 378–384.

2011

  • Ávila-Jiménez, M.L. & Coulson, S.J. (2011) A Holarctic biogeographical analysis of the Collembola (Arthropoda, Hexapoda) unravels recent post-glacial colonization patterns. Insects 2; 273-296
  • Gwiazdowicz  D.J. & Coulson S.J. (2011) High Arctic gamasid mites (Acari, Mesostigmata); community composition on Spitsbergen, Svalbard. Polar Research 30; 1-7.
  • Ávila-Jiménez M.L & Coulson S.J. (2011) Can snow depth predict the distribution of the high Arctic aphid Acyrthosiphon svalbardicum (Hemiptera: Aphididae) on Spitsbergen? BMC Ecology 11:25
  • Ávila-Jiménez M.L., Gwiazdowicz  D.J. & Coulson S.J. (2011) On the gamasid (Acari: Parasitiformes) mite fauna of Svalbard: a revised checklist of a high Arctic archipelago. Zootaxa  3091; 33–41.
  • Coulson S.J., Ávila-Jiménez M.L., Fjellberg A., Snazell R., & Gwiazdowicz D.J. (2011) On the Collembola, Araneae and Gamasida from the Kinnvika region of Nordaustlandet, Svalbard. Geografiska Annaler 93 (4); 253-257.
  • Gwiazdowicz  D.J., Teodorowicz E. & Coulson S.J. (2011) Redescription of Arctoseius haarlovi Lindquist, 1963 (Acari, Ascidae) from Spitsbergen, Svalbard. Entomologica Fennica. 22;140-148.
  • Gwiazdowicz  D.J., Teodorowicz E. & Coulson S.J. (2011) Redescription of Zercon solenites Haarløv, 1942 (Acari, Zerconidae) with a key to the Svalbard species of the genus Zercon. International Journal of Acarology  37(sup1);135-148.  

2010

  • Ávila-Jiménez, M. L., Coulson S.J , Solhøy T. & Sjöblom A. (2010) Overwintering of Arctic arthropods; the case of the invertebrate fauna of Svalbard now and in the future. Polar Research 29;127-137.
  • Gwiazdowicz D.J. & S.J. Coulson. (2010) Phoresis of the duetonymphs of Thinoseius spinosus (Acari, Eviphididae) on the calliphorid dipteran Protophormia terraenovae on the High Arctic island of Spitsbergen (Svalbard) including a key to deutonymphs of Thinoseius. International Journal of Acarology 36;233-236. 

2009

  • Coulson S.J. (2009) Association of the soil mite Diapterobates notatus (Acari, Oribatidae) with Cynomya mortuorum (Linnaeus, 1761) (Calliphoridae, Calliphorinae): implications for the dispersal of oribatid mites. International Journal of Acarology 35;175-177
  • Gwiazdowicz D.J, S.J.Coulson & M.L.Ávila-Jiménez.  (2009) First records of Zercon andrei Sellnick, 1958 and Zerconopsis muestairi (Schweizer, 1949) (Acari, Mesostigmata) from Bjørnøya. Norwegian Journal of Entomology 56, 117–119 
  • Coulson S.J., Monson F., Moe B. & Gabrielsen G.W. (2009) The invertebrate fauna of High Arctic seabird nests:  the microarthropod community inhabiting nests on Spitsbergen, Svalbard. Polar Biology 32;1041-1046.  
  • Coulson, S.J., Lorentzen, E., Strøm, H. Gabrielsen G.W. (2009) The parasitic tick Ixodes uriae (Acari: Ixodidae) on seabirds from Spitsbergen, Svalbard. Polar Research 28;399-402 

2008

  • Ávila-Jiménez, M. L., Fjellberg, A.  & Coulson, S.J. (2008) First record of Folsomia bisetosella Fjellberg, 2005 (Hexapoda: Collembola) from High Arctic islands. Norwegian Journal of Entomology  55, 129–130. 

2007

  • Coulson, S.J. 2007 The terrestrial and freshwater invertebrate fauna of the High Arctic archipelago of Svalbard. Zootaxa 1448; 41-58.  
  • Coulson, S.J. 2007 On the occurrence of Oryzaephilus mercator on Svalbard, Norway.  Norwegian Journal of Entomology. 54, 21-22.
  • White, D.M., Hodkinson I.D., Seelen S.J. & Coulson S.J. 2007. Characterization of soil carbon from a Svalbard glacier-retreat chronosequence using Pyrolysis-GC/MS analysis. Journal of Analytical and Applied Pyrolysis 78; 70-75. 

2004

  • Coulson, S.J. & Refseth, D. (2004) The terrestrial and freshwater invertebrate fauna of Svalbard (and Jan Mayen), pp57-122 in A catalogue of the terrestrial and marine animals of Svalbard (eds. Prestrud, P. Strøm H. & Goldman H.). Skrifter 201. Norwegian Polar Institute, Tromsø.
  • Cooper, E.J., Alsos, I.G., Hagen, D., Smith, F.M., Coulson, S.J. & Hodkinson, I.D. (2004) Plant recruitment in the Arctic: Seedbank and seeding emergence on Svalbard.  Journal of Vegetation Science 15; 115-124.
  • Glasser, N.F., Coulson, S.J., Hodkinson, I.D. & Webb, N.R. (2004) Photographic evidence of the return period of a Svalbard surge-type glacier: Pedersenbreen, Kongsfjord. Journal of Glaciology. 50(169); 307-308.
  • Hodkinson, I.D., Coulson, S. J. & Webb, N.R. (2004) Invertebrate community assembly along proglacial chronosequences in the high Arctic. Journal of Animal Ecology 73; 556-568
  • Hodkinson I.D. & Coulson S.J. (2004) Are high Arctic terrestrial food webs really that simple? - Bear Island revisited. Oikos 106; 427-431
  • Hodkinson, I.D., Bird, J.M. Cooper, E.J. & Coulson, S.J. (2004) The sexual morphs of the endemic Svalbard aphid Acyrthosiphon calvulus (Ossiannilsson), with notes on species biology. Norwegian Journal of Entomology 51; 131-135
  • Ims R.A.,  Leinaas, H.P. & Coulson, S.J.  (2004) Spatial and temporal variation in patch occupancy and population density in a model system of an arctic species assemblage of Collembola. Oikos 105; 89-100.
  • Hodkinson, I.D., Neerup Buhl, P., Coulson, S.J. & Webb N.R. (2004) First record of Dendrocerus bifoveatus (Kieffer, 1907)(Hymenoptera: Ceraphronoidea) from Svalbard. Norwegian Journal of Entomology 50; 105-106. 

2003

  • Hodkinson, I.D., Coulson, S. J. & Webb, N.R. (2003) Community assembly on proglacial chronosequences in the high Arctic: vegetation and soil development in north west Svalbard. Journal of Ecology 91; 651-653
  • Coulson, S.J., Hodkinson, I.D. Webb, N.R. & Convey, P. (2003) A high Arctic population of  Pyla fusca (Lepidoptera, Pyralidae) on Svalbard? Polar Biology 26; 283-285.
  • Coulson, S.J., Hodkinson, I.D. & Webb, N.R. (2003)  Microscale distribution patterns in high Arctic soil microarthropod communities: the influence of the vegetation mosaic. Ecography 26; 801-809. 
  • Coulson, S. J., Hodkinson, I.D., & Webb, N.R. (2003) Aerial dispersal of invertebrates over a High Arctic glacier foreland: Midtre Lovénbreen, Svalbard. Polar Biology 26; 530-537
  • Coulson, S. J., Hodkinson, I.D., Jussila R. & Webb, N.R. (2003) Synodites lineiger (Thomson, 1893) (Hymenoptera: Ichneumonidae) new to Svalbard, together with an updated list of the known ichneumonid species. Norwegian Journal of Entomology 50; 55-56  

2002

  • Coulson, S. J., Hodkinson, I.D., Webb, N.R. & Harrison, J. A.  (2002) Survival of saltwater immersion by terrestrial invertebrates. Implications for the colonisation of Arctic islands. Functional. Ecology 16; 353-356  
  • Coulson, S. J., Hodkinson, I.D., Webb, N.R.. Mikkola, K., Harrison, J. A. & Pedgley, D.  (2002) Aerial colonisation of high Arctic islands by invertebrates: the Diamondback Moth, Plutella xylostella (Lepidoptera: Yponomeutidae) as a potential indicator species. Diversity and Distributions 8; 327-334.
  • Hodkinson I.D., Coulson S.J. & Bird J.M. (2002)  Discovery of a rare alate morph of Acyrthosiphon svalbardicum Heikinheimo (Homoptera; Aphididae), a hitherto ‘apterous’ aphid species: description and implications for species ecology. Norwegian Journal of Entomology 49; 77-80.
  • Hodkinson, I.D Webb, N.R. & Coulson, S. J. (2002) Primary community assembly on land: why are the predators and scavengers always there first? Are we missing a vital stage? Journal of Ecology 90; 569-577.  

2001

  • Coulson, S. J., Hodkinson, I.D., Jussila, R., Webb, N.R. & Harrison, J. A. (2001) Atractodes pusillus Förster, 1876 and Stenomacrus groenlandicus Jussila, 1996 (Hymenoptera: Ichneumonidae), new records to Svalbard, Norway.  Norwegian Journal of Entomology 48; 328
  • Hodkinson I.D. Coulson, S.J. Harrison J. & Webb N.R. (2001) What a wonderful web they weave: spiders, nutrient capture in early ecosystem development in the High Arctic - some counter intuitive ideas on community assembly. Oikos 95; 349-352 

2000

  • Coulson, S.J. & Birkemoe, T. (2000) Long term cold tolerance in Arctic invertebrates: recovery after four years at below -20°C. Canadian Journal of Zoology 78; 2055-2058.
  • Coulson, S.J., (2000) A review of the terrestrial and freshwater invertebrate fauna of Svalbard, a High Arctic island group. Norwegian  Journal of Entomology 47; 41-63. 
  • Coulson, S.J., Leinaas, H.P., Ims R.A. & Søvik G. (2000) Experimental manipulation of the  winter surface ice layer: the effects on a High Arctic soil microarthropod community. Ecography 23; 299-314
  • Birkemoe, T., Coulson, S.J. & Sømme, L. (2000) Life cycles and population dynamics of enchytraeids (Oligochaeta) from the high Arctic. Canadian Journal of Zoology 78; 2079-2086. 

1998

  • Hodkinson I.D., Webb N.R., Bale J.S., Coulson S.J. & Strathdee A.T. (1998) Global change and high arctic ecosystems: conclusions and predictions from experiments with terrestrial invertebrates on Spitsbergen. Arctic and Alpine Research, 30; 306-313.
  • Webb, N.R.,  Coulson, S.J., Hodkinson, I.D., Block, W.,  Bale, J.S. & Strathdee, A.T.  (1998) The effects of experimental temperature elevation on populations of cryptostigmatic mites in high Arctic soils. Pedobiologia. 42; 298-308.  

1996

  • Coulson S.J. & Bale J.S. (1996) Supercooling and survival of the beech weevil Rhynchaenus fagi L. (Coleoptera: Curculionidae). Journal of Insect Physiology 42; 617-623 
  • Coulson S.J., Hodkinson I.D., Webb N.R., Block W., Bale J.S., Strathdee A.T., Worland M.R. & Wooley C. (1996) Effects of experimental temperature elevation on high-arctic soil microarthropod populations. Polar Biology 16; 147-153.
  • Hodkinson I.D., Coulson S.J. Webb N.R. & Block W. (1996) Can Arctic soil microarthropods survive elevated summer temperatures? Functional Ecology 10; 314-321.
  • Hodkinson I.D., Coulson S.J., Webb N.R., Block W., Strathdee A.T., Bale J.S., & Worland M.R. (1996) Temperature and the biomass of flying midges (Diptera: Chironomidae) in a high Arctic ecosystem. Oikos 75; 241-248. 

1995

  • Coulson S.J., Hodkinson I.D., Block W., Webb N.R. & Worland M.R. (1995) Low summer temperatures: a potential mortality factor for high Arctic soil micro-arthropods? Journal of Insect Physiology 41; 783-792.
  • Coulson S.J., Hodkinson I.D., Strathdee A.T., Block W., Webb N.R., Bale J.S. & Worland M.R. (1995) Thermal environments of Arctic soil organisms during winter. Arctic and Alpine Research 27; 365-371.
  • Strathdee A.T., Bale J.S., Strathdee F.C., Block W., Coulson S.J., Hodkinson I.D. & Webb N.R. (1995) Climatic severity and the response to warming of Arctic aphids. Global Change Biology 1; 23-28. 

1994

  • Hodkinson I.D., Healey V. & Coulson S.J. (1994)  Moisture relationships of the high arctic collembolan Onychiurus arcticus. Physiological Entomology 19; 109-114.
  • Block W., Webb N.R., Coulson S.J., Hodkinson I.D. (1994) Thermal adaptation in a high arctic collembolan Onychiurus arcticus. Journal of Insect Physiology 40; 715-722.  

1993

  • Coulson S.J., Hodkinson I.D., Strathdee A.T., Bale J.S., Block W., Worland M.R. & Webb N.R. (1993) Simulated climate change: the interaction between vegetation type and microhabitat temperatures at Ny Ålesund, Svalbard. Polar Biology 13; 67-70.
  • Hodkinson I.D., Coulson S.J., Webb N.R., Block W., Strathdee A.T. & Bale J.S. (1993) Feeding studies on Onychiurus arcticus (Tullberg) Collembola: Onychiuridae) on West Spitsbergen. Polar Biology 14; 17-19.
  • Strathdee A.T., Bale J.S., Hodkinson I.D., Block W., Webb N.R. & Coulson S.J. (1993) Identification of three previously unknown morphs of Acyrthosiphon svalbardicum Heikinheimo (Hemiptera: Aphididae) on Spitsbergen. Entomologica Scandinavica 24; 43-47.
  • Strathdee A.T., Bale J.S., Block W.C., Webb N.R., Hodkinson I.D. & Coulson S.J. (1993) Extreme adaptive life-cycle in a high arctic aphid, Acyrthosiphon svalbardicum. Ecological Entomology 18; 254-258.
  • Strathdee, A.T., Bale, J.S., Block, W., Coulson, S.J., Hodkinson, I.D. & Webb, N.R. (1993) Effects of temperature elevation on a field population of Acyrthosiphon svalbardicum (Hemiptera: Aphididae) on Spitsbergen. Oecologia 96; 457-465. 

1992

  • Coulson S.J. & Bale J.S. (1992) Effect of rapid cold hardening on reproduction and survival of offspring in the housefly Musca domestica. Journal of Insect Physiology 38; 421-424. 
  • Coulson S.J., Fisher J. & Bale J.S. (1992) A 31P NMR investigation of the energy charge of the housefly Musca domestica (Diptera: Muscidae). During rapid cold hardening and cold shock. Cryo-Letters 13; 183-192. 

1991

  • Coulson S.J. & Bale J.S. (1991) Anoxia induces rapid cold hardening in the housefly Musca domestica (Diptera: Muscidae). Journal of Insect Physiology 37; 497-501. 

1990

  • Coulson S.J. & Bale J.S. (1990) Characterisation and limitations of the rapid cold hardening response in the housefly Musca domestica (Diptera: Muscidae). Journal of Insect Physiology 36; 201-211.

Back to previous page

 

 


Oribatid mite, Diapterobates notatus ((Thorell 1871), ca. 0.5mm in length

Head of the collembolan Ceratophysella longispina (Tullberg 1876)

 

 

Search the UNIS web
Things of interest

APPLY ONLINE TO UNIS
Find information about how to become a student at UNIS

UNIS STUDENT MOVIE
SIOS
Svalbard Integrated Arctic Earth Observing System
UNIS CO2 LAB

TOTAL SOLAR ECLIPSE
20 MARCH 2015

KJELL HENRIKSEN OBSERVATORY
(Aurora station)

SPEAR RADAR

BIRKELAND CENTRE for SPACE SCIENCE

WEATHER
Realtime weather data and downloadable historical data.

VACANT POSITIONS

ANNUAL REPORTS

360° PANORAMA CAMERA
Picture from UNIS right now

 

Like us on Facebook

The University Centre in Svalbard | Pb. 156 | 9171 Longyearbyen, Norway | Tel: +47 79 02 33 00 - Fax: +47 79 02 33 01| Org. 985 204 454 | post@unis.no